Now, it cannot be held that the present races of animals differ from the ancient races merely by modifications produced by local circ.u.mstances and change of climate--for if species gradually changed, we must find traces of these gradual modifications, and between the palaeotheria and the present species we should have discovered some intermediate formation; but to the present time none of these have appeared.
Why have not the bowels of the earth preserved the monuments of so remarkable a genealogy unless it be that the species of former ages were as constant as our own, or at least because the catastrophe that destroyed them had not left them time to give evidence of the changes?
Further, an examination of animals shows that though their superficial characteristics, such as colour and size, are changeable, yet their more radical characteristics do not change. Even the artificial breeding of domestic animals can produce only a limited degree of variation. The maximum variation known at the present time in the animal kingdom is seen in dogs, but in all the varieties the relations of the bones remain the same and the shape of the teeth undergoes no palpable change.
I know that some naturalists rely much on the thousands of ages which they can acc.u.mulate with a stroke of the pen; but there is nothing which proves that time will effect any more than climate and a state of domestication. I have endeavoured to collect the most ancient doc.u.ments of the forms of animals. I have examined the engravings of animals including birds on the numerous columns brought from Egypt to Rome. M.
Saint Hilaire collected all the mummies of animals he could obtain in Egypt--cats, ibises, birds of prey, dogs, monkeys, crocodiles, etc.--and we cannot find any more difference between them and those of the present day than between human mummies of that date and skeletons of the present day.
There is nothing, then, in known facts which can support the opinion that the new genera discovered among fossils--the palaeotheria, anoplotheria, megalonyces, mastodontes, pterodactyli, ichthyosauri, etc.--could have been the sources of any animals now existing, which would differ only by the influence of time or climate.
As yet no human bones have been discovered in the regular layers of the surface of the earth, so that man probably did not exist in the countries where fossil bones are found at the epoch of the revolutions which buried these bones, for there cannot be a.s.signed any reason why mankind should have escaped such overwhelming catastrophes, or why human remains should not be discovered. Man _may_ have inhabited some confined tract of country which escaped the catastrophe, but his establishment in the countries where the fossil remains of land animals are found--that is to say, in the greatest part of Europe, Asia, and America--is necessarily posterior not only to the revolutions which covered these bones, but even to those which have laid open the strata which envelop them; whence it is clear that we can draw neither from the bones themselves nor from the rocks which cover them any argument in favour of the antiquity of the human species in these different countries. On the contrary, in closely examining what has taken place on the surface of the globe, since it was left dry for the last time, we clearly see that the last revolution, and consequently the establishment of present society, cannot be very ancient. An examination of the amount of alluvial matter deposited by rivers, of the progress of downs, and of other changes on the surface of the earth, informs us clearly that the present state of things did not commence at a very remote period.
The history of nations confirms the testimony of the fossils and of the rocks. The chronology of none of the nations of the West can be traced unbroken farther back than 3,000 years. The Pentateuch, the most ancient doc.u.ment the world possesses, and all subsequent writings allude to a universal deluge, and the Pentateuch and Vedas and Chou-king date this catastrophe as not more than 5,400 years before our time. Is it possible that mere chance gave a result so striking as to make the traditional origin of the a.s.syrian, Indian, and Chinese monarchies agree in being as remote as 4,000 or 5,000 years back? Would the ideas of nations with so little inter-communication, whose language, religion, and laws have nothing in common, agree on this point if they were not founded on truth? Even the American Indians have their Noah or Deucalion, like the Indians, Babylonians, and Greeks.
It may be said that the long existence of ancient nations is attested by their progress in astronomy. But this progress has been much exaggerated. But what would this astronomy prove even if it were more perfect? Have we calculated the progress which a science would make in the bosom of nations which had no other? If among the mult.i.tude of persons solely occupied with astronomy, even then, all that these people knew might have been discovered in a few centuries, when only 300 years intervened between Copernicus and Laplace.
Again, it has been pretended that the zodiacal figures on ancient temples give proof of a remote antiquity; but the question is very complicated, and there are as many opinions as writers, and certainly no conclusions against the newness of continents and nations can be based on such evidence. The zodiac itself has been considered a proof of antiquity, but the arguments brought forward are undoubtedly unsound.
Even if these various astronomical proofs were as certain as they are unconvincing, what conclusion could we draw against the great catastrophe so indisputably demonstrated? We should only have the right to conclude that astronomy was among the sciences preserved by those persons whom the catastrophe spared.
In conclusion, if there be anything determined in geology, it is that the surface of our globe has been subjected to a revolution within 5,000 years, and that this revolution buried the countries formerly inhabited by man and modern animals, and left the bottom of the former sea dry as a habitation for the few individuals it spared. Consequently, our present human societies have arisen since this catastrophe.
But the countries now inhabited had been inhabited before, as fossils show, by animals, if not by mankind, and had been overwhelmed by a previous deluge; and, indeed, judging by the different orders of animal fossils we find, they had perhaps undergone two or three irruptions of the sea.
CHARLES DARWIN
The Origin of Species
Charles Robert Darwin was born at Shrewsbury, England, Feb. 12, 1809, of a family distinguished on both sides. Abandoning medicine for natural history, he joined H.M.S. Beagle in 1831 on the five years" voyage, which he described in "The Voyage of the Beagle,"
and to which he refers in the introduction to his masterpiece. The "Origin of Species" containing, in the idea of natural selection, the distinctive contribution of Darwin to the theory of organic evolution, was published in November, 1859. In only one brief sentence did he there allude to man, but twelve years later he published the "Descent of Man," in which the principles of the earlier volume found their logical outcome. In other works Darwin added vastly to our knowledge of coral reefs, organic variation, earthworms, and the comparative expression of the emotions in man and animals. Darwin died in ignorance of the work upon variation done by his great contemporary, Gregor Mendel, whose work was rediscovered in 1900. "Mendelism" necessitates much modification of Darwin"s work, which, however, remains the maker of the greatest epoch in the study of life and the most important contribution to that study ever made. Its immortal author died on April 19, 1882, and was buried in Westminster Abbey.
_I.--Creation or Evolution?_
When on board H.M.S. Beagle as naturalist, I was much struck with certain facts in the distribution of the organic beings inhabiting South America, and in the geographical relations of the present to the past inhabitants of that continent. These facts, as will be seen in the latter chapters of this volume, seemed to throw some light on the origin of species--that mystery of mysteries, as it has been called by one of our greatest philosophers. On my return home, in 1837, it occurred to me that something might perhaps be made out on this question by patiently acc.u.mulating and reflecting on all sorts of facts which could possibly have any bearing on it. After five years" work, I allowed myself to speculate on the subject, and drew up some short notes; these I enlarged in 1844 into a sketch of the conclusions which then seemed to me probable. From that period to the present day I have steadily pursued the same object. I hope that I may be excused for entering on these personal details, as I give them to show that I have not been hasty in coming to a decision.
In considering the origin of species, it is quite conceivable that a naturalist, reflecting on the mutual affinities of organic beings, on their embryological relations, their geographical distribution, geological succession, and other such facts, might come to the conclusion that species had not been independently created, but had descended, like varieties, from other species. Nevertheless, such a conclusion, even if well founded, would be unsatisfactory, until it could be shown how the innumerable species inhabiting this world have been modified so as to acquire that perfection of structure and co-adaptation which justly excites our admiration.
Naturalists continually refer to external conditions, such as climate, food, etc., as the only possible cause of variation. In one limited sense, as we shall hereafter see, this may be true; but it is preposterous to attribute to mere external conditions the structure, for instance, of the woodp.e.c.k.e.r, with its feet, tail, beak, and tongue, so admirably adapted to catch insects under the bark of trees. In the case of the mistletoe, which draws its nourishment from certain trees, which has seeds that must be transported by certain birds, and which has flowers with separate s.e.xes absolutely requiring the agency of certain insects to bring pollen from one flower to the other, it is equally preposterous to account for the structure of the parasite, with its relations to several distinct organic beings, by the effects of external conditions, or of habit, or of the volition of the plant itself.
It is, therefore, of the highest importance to gain a clear insight into the means of modification and co-adaptation. At the beginning of my observations it seemed to me probable that a careful study of domesticated animals and of cultivated plants would offer the best chance of making out this obscure problem. Nor have I been disappointed; in this and in all other perplexing cases I have invariably found that our knowledge, imperfect though it be, of variation under domestication, afforded the best and safest clue. I may venture to express my conviction of the high value of such studies, although they have been very commonly neglected by naturalists.
Although much remains obscure, and will long remain obscure, I can entertain no doubt, after the most deliberate study and dispa.s.sionate judgment of which I am capable, that the view which most naturalists until recently entertained, and which I formerly entertained--namely, that each species has been independently created--is erroneous. I am fully convinced that species are not immutable, but that those belonging to what are called the same genera are lineal descendants of some other and generally extinct species, in the same manner as the acknowledged varieties of any one species are the descendants of that species.
Furthermore, I am also convinced that Natural Selection has been the most important, but not the exclusive, means of modification.
_II.--Variation and Selection_
All living beings vary more or less from one another, and though variations which are not inherited are unimportant for us, the number and diversity of inheritable deviations of structure, both those of slight and those of considerable physiological importance, are endless.
No breeder doubts how strong is the tendency to inheritance; that like produces like is his fundamental belief. Doubts have been thrown on this principle only by theoretical writers. When any deviation of structure often appears, and we see it in the father and child, we cannot tell whether it may not be due to the same cause having acted on both; but when amongst individuals, apparently exposed to the same conditions, any very rare deviation, due to some extraordinary combination of circ.u.mstances, appears in the parent--say, once amongst several million individuals--and it re-appears in the child, the mere doctrine of chances almost compels us to attribute its reappearance to inheritance.
Everyone must have heard of cases of albinism, p.r.i.c.kly skin, hairy bodies, etc., appearing in members of the same family. If strange and rare deviations of structure are really inherited, less strange and commoner deviations may be freely admitted to be inheritable. Perhaps the correct way of viewing the whole subject would be to look at the inheritance of every character whatever as the rule, and non-inheritance as the anomaly.
The laws governing inheritance are for the most part unknown. No one can say why the same peculiarity in different individuals of the same species, or in different species, is sometimes inherited and sometimes not so; why the child often reverts in certain characters to its grandfather or grandmother, or more remote ancestor; why a peculiarity is often transmitted from one s.e.x to both s.e.xes, or to one s.e.x alone, more commonly but not exclusively to the like s.e.x.
The fact of heredity being given, we have evidence derived from human practice as to the influence of selection. There are large numbers of domesticated races of animals and plants admirably suited in various ways to man"s use or fancy--adapted to the environment of which his need and inclination are the most essential const.i.tuents. We cannot suppose that all the breeds were suddenly produced as perfect and as useful as we now see them; indeed, in many cases, we know that this has not been their history. The key is man"s power of acc.u.mulative selection. Nature gives successive variations; man adds them up in certain directions useful to him. In this sense he may be said to have made for himself useful breeds.
The great power of this principle of selection is not hypothetical. It is certain that several of our eminent breeders have, even within a single lifetime, modified to a large extent their breeds of cattle and sheep. What English breeders have actually effected is proved by the enormous prices given for animals with a good pedigree; and these have been exported to almost every quarter of the world. The same principles are followed by horticulturists, and we see an astonishing improvement in many florists" flowers, when the flowers of the present day are compared with drawings made only twenty or thirty years ago.
The practice of selection is far from being a modern discovery. The principle of selection I find distinctly given in an ancient Chinese encyclopaedia. Explicit rules are laid down by some of the Roman cla.s.sical writers. It is clear that the breeding of domestic animals was carefully attended to in ancient times, and is now attended to by the lowest savages. It would, indeed, have been a strange fact had attention not been paid to breeding, for the inheritance of good and bad qualities is so obvious.
Study of the origin of our domestic races of animals and plants leads to the following conclusions. Changed conditions of life are of the highest possible importance in causing variability, both by acting directly on the organisation, and indirectly by affecting the reproductive system.
Spontaneous variation of unknown origin plays its part. Some, perhaps a great, effect may be attributed to the increased use or disuse of parts.
The final result is thus rendered infinitely complex. In some cases the intercrossing of aboriginally distinct species appears to have played an important part in the origin of our breeds. When several breeds have once been formed in any country, their occasional intercrossing, with the aid of selection, has, no doubt, largely aided in the formation of new sub-breeds; but the importance of crossing has been much exaggerated, both in regard to animals and to those plants which are propagated by seed. Over all these causes of change, the acc.u.mulative action of selection, whether applied methodically and quickly, or unconsciously and slowly, but more efficiently, seems to have been the predominant power.
_III.--Variation Under Nature_
Before applying these principles to organic beings in a state of nature, we must ascertain whether these latter are subject to any variation. We find variation everywhere. Individual differences, though of small interest to the systematist, are of the highest importance for us, for they are often inherited; and they thus afford materials for natural selection to act and acc.u.mulate, in the same manner as man acc.u.mulates in any given direction individual differences in his domesticated productions. Further, what we call varieties cannot really be distinguished from species in the long run, a fact which we can clearly understand if species once existed as varieties, and thus originated.
But the facts are utterly inexplicable if species are independent creations.
How have all the exquisite adaptations of one part of the body to another part, and to the conditions of life, and of one organic being to another being, been perfected? For everywhere we find these beautiful adaptations.
The answer is to be found in the struggle for life. Owing to this struggle, variations, however slight, and from whatever cause proceeding, if they be in any degree profitable to the individuals of a species in their infinitely complex relations to other organic beings and to their physical conditions of life, will tend to the preservation of such individuals, and will generally be inherited by the offspring.
The offspring, also, will thus have a better chance of surviving, for, of the many individuals of any species which are periodically born, but a small number can survive. I have called this principle, by which each slight variation, if useful, is preserved, by the term Natural Selection, in order to mark its relation to man"s power of selection.
But the expression, often used by Mr. Herbert Spencer, of the Survival of the Fittest, is more accurate.
We have seen that man, by selection, can certainly produce great results, and can adapt organic beings to his own uses, through the acc.u.mulation of slight but useful variations given to him by the hand of Nature. Natural Selection is a power incessantly ready for action, and is as immeasurably superior to man"s feeble efforts as the works of Nature are to those of Art.
All organic beings are exposed to severe compet.i.tion. Nothing is easier than to admit in words the truth of the universal struggle for life, or more difficult--at least, I have found it so--than constantly to bear this conclusion in mind. Yet, unless it be thoroughly engrained in the mind, the whole economy of Nature, with every fact of distribution, rarity, abundance, extinction, and variation, will be dimly seen or quite misunderstood. We behold the face of Nature bright with gladness; we often see superabundance of food. We do not see, or we forget, that the birds which are idly singing round us mostly live on insects or seeds, and are thus constantly destroying life; or we forget how largely these songsters, or their eggs, or their nestlings, are destroyed by birds or beasts of prey. We do not always bear in mind that, though food may be superabundant, it is not so at all seasons of each recurring year.
A struggle for existence, the term being used in a large, general, and metaphorical sense, inevitably follows from the high rate at which all organic beings tend to increase.
Every being, which during its natural lifetime produces several eggs or seeds, must suffer destruction during some period of its life, and during some season or occasional year; otherwise, on the principle of geometrical increase, its numbers would quickly become so inordinately great that no country could support the product. Hence, as more individuals are produced than can possibly survive, there must in every case be a struggle for existence, either one individual with another of the same species, or with the individuals of distinct species, or with the physical conditions of life. It is the doctrine of Malthus applied with manifold force to the whole animal and vegetable kingdoms; for in this case there can be no artificial increase of food, and no prudential restraint from marriage. Although some species may be now increasing, more or less rapidly, in numbers, all cannot do so, for the world would not hold them.
There is no exception to the rule that every organic being naturally increases at so high a rate that, if not destroyed, the earth would soon be covered by the progeny of a single pair. Even slow-breeding man has doubled in twenty-five years, and at this rate, in less than a thousand years, there would literally not be standing-room for his progeny.
Linnaeus has calculated that if an annual plant produced only two seeds--and there is no plant so unproductive as this--and their seedlings next year produced two, and so on, then in twenty years there would be a million plants. The elephant is reckoned the slowest breeder of all known animals, and I have taken some pains to estimate its probable minimum rate of natural increase. It will be safest to a.s.sume that it begins breeding when thirty years old, and goes on breeding until ninety years old, bringing forth six young in the interval, and surviving till one hundred years old. If this be so, after a period of from 740 to 750 years there would be nearly nineteen million elephants alive, descended from the first pair.
The causes which check the natural tendency of each species to increase are most obscure. Eggs or very young animals seem generally to suffer most, but this is not invariably the case. With plants there is a vast destruction of seeds. The amount of food for each species of course gives the extreme limit to which each can increase; but very frequently it is not the obtaining food, but the serving as prey to other animals, which determines the average number of a species. Climate is important, and periodical seasons of extreme cold or drought seem to be the most effective of all checks.
The relations of all animals and plants to each other in the struggle for existence are most complex, and often unexpected. Battle within battle must be continually recurring with varying success; and yet in the long run the forces are so nicely balanced that the face of Nature remains for long periods of time uniform, though a.s.suredly the merest trifle would give the victory to one organic being over another.
Nevertheless, so profound is our ignorance, and so high our presumption, that we marvel when we hear of the extinction of an organic being; and as we do not see the cause, we invoke cataclysms to desolate the world, or invent laws on the duration of the forms of life!